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Participants used a combination of boxing and wrestling techniques, but also others, such as kicking and holds, locks and chokes on the ground. The only things not acceptable were biting and gouging out the opponent's eyes. 
By the time of the Peloponnesian War, the majority of Greek poleis had a pankration champion representing them at the Olympic Games; in 428 BCE, the pankratists participating in the event included Orion and Orestes, while Rhodes was represented by the reigning champion Dorieus, and Sparta was represented by the misthios Kassandra, who had to take the place of Sparta's Testikles following his untimely death.