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Maegesters were high-ranking Order of the Ancients members who led a specific branch of the organization. They answered directly to the Grand Maegester. In the 870s, all five active Maegesters in England were hunted down and killed by Eivor Varinsdottir. 
Behind the scenes
The title of Maegester uses an alternate spelling of the Latin word Magister, which means "teacher" or "master", and serves as a linguistic prelude to the current title of Master Templar used to denote a Templar with sufficient authority to lead a Rite if a Grand Master is not present.