Colosseum Vault

The Colosseum Vault was a chamber created by Those Who Came Before located beneath the Santa Maria Aracoeli. The vault was created to store the Apple of Eden on an erected pedestal at the center of a large chamber and also contained what remained of the goddess, Juno.

To enter the Vault's central chamber, one must speak aloud the number "72." After this, the tall pedestal can only accessed by jumping on a series of thin platforms and levers that appear to activate when a person with certain DNA, such as Desmond Miles or Ezio Auditore, touches them, most likely because of being descendants of Those Who Came Before, and would therefore be allowed access to their technology. After activating all platforms, a set of stairs rise to form an easy path to the Piece of Eden.

Visitors
The earliest known visitor of the Vault was Marcus Junius Brutus, as evidenced by the 4th Scroll of Romulus. Brutus also made a drawing of the Vault, as well as the pedestal holding the Piece of Eden.

The next known visitor was Ezio Auditore, who knew the password to enter (possibly through the use of the Apple of Eden). He then placed the Apple of Eden on the pedestal and left.

The latest known visitors were Desmond Miles, Lucy Stillman, Shaun Hastings and Rebecca Crane; who figured out the password from the writing Ezio left in the Villa Auditore. Entering into the vault, Desmond met with the goddess Juno, who spoke to him 4 times in his attempt to aquire the Apple of Eden, and out of all the Assassin's present was the only one able to see and hear her. After Desmond activated the stairs and walked up to the Apple of Eden, symbols appeared around all of them. Upon touching the Apple, Desmond lost control of his body as Juno took over his actions, and forced to stab Lucy in the abdomen before collapsing on the floor.

Trivia

 * Shaun remarks on two of the symbols that appear when the Apple is activated, the Phrygian cap and the Masonic Eye. He says "Now those two come together in only one place-" before being frozen.
 * The above symbols only appear together on the Revolutionary France 'Declaration of the Rights of Man and of the Citizen', written in August 1798.
 * Of the symbols that had appeared the Abstergo Logo does appear quite frequently.

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